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Doctoral Thesis
DOI
10.11606/T.41.2009.tde-16122009-182801
Document
Author
Full name
Cibele Bragagnolo
E-mail
Institute/School/College
Knowledge Area
Date of Defense
Published
São Paulo, 2009
Supervisor
Committee
Rocha, Ricardo Pinto da (President)
Carvalho, Marcelo Rodrigues de
Hara, Marcos Ryotaro
Kury, Adriano Brilhante
Nihei, Silvio Shigueo
Title in Portuguese
Revisão sistemática de Promitobates Roewer 1913 e análise filogenética de Mitobatinae Simon1879 (Arachnida, Opiliones, Gonyleptidae)
Keywords in Portuguese
Gonyleptidae
Opilioes
Sistemática
Abstract in Portuguese
As 10 especies nominais de Promitobates foram utilizadas na analise cladistica da subfamilia Mitobatinae. A especie P. ornatus (Mello-Leitao, 1922), amplamente distribuida e polimorfica, foi decomposta em quatro unidades monomorficas e sua taxonomia testada durante a analise cladistica. Outras duas especies que apresentaram polimorfismos (P. hatschbachi H. Soares, 1945 e P. viridigranulatus Soares & Soares, 1946) tambem foram separadas em duas unidades distintas. A analise cladistica foi efetuada com mais 20 especies de Mitobatinae, representando os 11 generos que formam a subfamilia e mais cinco representantes de Gonyleptidae. A matriz e composta por 75 caracteres, sendo 19 pertencentes a genitalia masculina; 27 as estruturas morfologicas externas presentes no escudo dorsal, tergitos livres e pedipalpos; 20 referentes as estruturas presentes nas pernas dos machos e 10 de coloracao. Duas arvores igualmente parcimoniosas foram encontradas (L=257, C.I=0.38; R.I=0.72). Mitobatinae e monofiletico, sustentado por cinco sinapomorfias nao ambiguas e grupo-irmao de Despirus parvulus (Roewer, 1927). Mitobatinae e composto por dois grandes clados; [1] um grupo que possui corpo grosseiramente retangular e machos e femeas com a coxa e o trocanter IV inermes, ou com apofises muito reduzidas, incluindo os generos: Ischnotherus, Encheiridium, Metamitobates, Ruschia, Mitobatula e Mitobates e [2] um segundo grupo que apresenta corpo grosseiramente piriforme e dimorfismo sexual na coxa e trocanter IV, com os machos apresentando principalmente forte armacao externa na coxa e apofises no trocanter, incluindo os generos: Discocyrtoides, Longiperna, Neoancistrotus e Promitobates. O genero Promitobates Roewer 1913 e revisado e uma classificacao e proposta baseada nos resultados da analise cladistica. O genero era composto por 10 especies nominais, destas, quatro sao consideradas validas: P. bellus (B. Soares, 1945); P. hatschbachi H. Soares, 1945; P. ornatus (Mello-Leitao, 1922) e P. viridigranulatus (Soares & Soares, 1946). Tres sinonimias sao propostas: P. hauseri (Sylhavy, 1979) e considerado sinonimo de P. viridigranulatus, P. margaritatus Roewer, 1931, sinonimo de P. ornatus e P. mendax H. Soares, 1945, sinonimo de P. hatschbachi. P. granulosissimus Mello-Leitao, 1932 e P. hexacanthus Koch, 1839 sao consideradas species inquirendae. Com base nos resultados da analise de P. ornatus separado em unidades distintas na analise cladistica, P. intermedius e P. nigripes sao revalidadas e, juntamente com P. ornatus, abrigam as quatro unidades monomorficas do complexo P. ornatus. Quatro especies novas sao descritas: P. sp.n1 (localidade-tipo: Ano Bom, Santa Catarina); P. sp. n2 (localidade-tipo: Reserva Morro Grande, Sao Paulo); P. sp.n3 (Localidade-tipo: Ribeirao Pires, Sao Paulo) e P. sp.n4 (Localidade-tipo: Jacarepagua, Rio de Janeiro).
Title in English
Systematic Review of the genus Promitobates Roewer, 1913 and cladistic analysis of Mitobatinae Simon 1879 (Arachnida, Opiliones, Gonyleptidae)
Keywords in English
Gonyleptidae
Harvestmen
Systematic
Abstract in English
A cladistic analysis of the subfamily Mitobatinae is presented. All 10 of the currently valid species of the genus Promitobates Roewer, 1913 were taken into account. Promitobates ornatus (Mello-Leitao, 1922), a polymorphic and widespread species, was initially broken into four sub-units and its taxonomy addressed with the results of the cladistic analysis. The other two polymorphic species, P. hatschbachi H. Soares, 1945 and P. viridigranulatus Soares & Soares, 1946, were also broken into two sub-units. In addition to these species, the cladistic analysis was performed with 20 other Mitobatinae species, representing the 11 genera of the subfamily, and 5 species of other Gonyleptidae genera . The character matrix comprised 75 characters: 19 from male genitalia, 27 from the general external morphology, 20 from male legs and 10 from coloration. Two equally parsimonious trees were obtained (L=257, C.I=0.38; R.I=0.72). Mitobatinae arose as a monophyletic group, sister to Despirus parvulus (Roewer, 1927). The subfamily was divided into two major groups: [1] one that possesses the body roughly rectangular and males and females with coxa and trochanter IV without large apophysis, including the genera: Ischnotherus Kury, 1991, Encheiridium Kury, 2003, Metamitobates Roewer, 1913, Ruschia Mello-Leitao, 1940, Mitobatula Roewer, 1931 and Mitobates Sundevall, 1833 and [2] one that possesses the body roughly piriform and an evident sexual dimorphism of coxa and trochanter IV, with males with a large apophysis on coxa and tubercles on trochanter IV, including the genera: Discocyrtoides Mello-Leitao, 1923, Longiperna Roewer, 1929, Neoancistrotus Mello-Leitao, 1927 and Promitobates. A taxonomic review of the genus Promitobates Roewer 1913 and a new classification, supported by the cladistc analysis results, are presented . Promitobates comprised 10 described species of which four are considered valid: P. bellus (B. Soares, 1945); P. hatschbachi H. Soares, 1945; P. ornatus (Mello-Leitao, 1922) and P. viridigranulatus (Soares & Soares, 1946). The following species are synonymized: P. hauseri (Sylhavy, 1979), considered a senior synonym of P. viridigranulatus (Soares & Soares, 1946); P. margaritatus Roewer, 1931 considered a senior synonym of P. ornatus (Mello-Leitao, 1922) and P. mendax H. Soares, 1945, considered a senior synonym of P. hatschbachi H. Soares, 1945. Promitobates granulosissimus Mello-Leitao, 1932 and P. hexacanthus Koch, 1839 are considered species inquirendae. Based on the results of the cladistc analysis, with P. ornatus separated in four subunits, P. intermedius (Mello-Leitao 1935) and P. nigripes (Mello-Leitao, 1935) are revalidated. Four new species are described: P. sp. n1 (type-locality: Ano Bom, Santa Catarina); P. sp. n2 (type-locality: Reserva Morro Grande, Sao Paulo); P. sp.n3 (type-locality: Ribeirao Pires, Sao Paulo) and P. sp.n4 (typelocality: Jacarepagua, Rio de Janeiro).
 
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Publishing Date
2010-03-24
 
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